Slotting in between the XC and the Pro versions, the Trail provides a middle ground of protection. The trail insert features a series of little notches around the edges, which serve two main purposes: provide a little bit more "give" as you transition onto the side of the tire, to give a slightly softer feel to the tire in turns (the Pro is notably stiffer and can make itself felt as you transition onto the side knobs). The notches also provide extra channels for sealant to move around.
Credit
Vital MTB / Johan Hjord
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PIT BITS - Sea Otter 2023 Day 2